Published Papers


Click on Highlighted Names for an abstract or .pdf of each paper or chapter.

Calabrese, A. and S.C. Weeks. 2022. Testing a behavioral model for the maintenance of androdioecy as a result of sexual conflict in the clam shrimp Eulimnadia dahli. Ethology 128:331-338

Hethke, M., S.C. Weeks, V. Schöttle, and D.C. Rogers. 2021. Preliminary study of temperature effects on size and shape in the modern spinicaudatan Eulimnadia texana (Crustacea: Branchiopoda). Zoological Studies 60. DOI: 2 10.6620/zs.2021.60-02

Astrop, T.I., L. Park-Boush and S.C. Weeks. 2020. Testing Weissman’s lineage selection model for the maintenance of sex: The evolutionary dynamics of clam shrimp reproduction over geologic time. Zoological Studies 59:16.

Hethke, M. and S.C. Weeks. 2020. Population density effects on carapace growth in clam shrimp: Implications for palaeontological studies. Zoological Studies 59:34.

Weeks, S.C. 2020. Effects of dietary restriction on lifespan, growth and reproduction of the clam shrimp Eulimnadia texana. Hydrobiologia 14:3067-3076.

Hethke, M., F.T. Fürsich, B. Jiang, B. Wang, P. Chellouche and S.C. Weeks. 2019. Ecological stasis in Spinicaudata (Crustacea: Branchiopoda)? – Early cretaceous clam shrimp of the Yixian formation of NE China occupied a broader realized ecological niche than extant members of the group. Palaeontology 62:483-513.

Calabrese, A. and S.C. Weeks. 2018. Are all-hermaphroditic populations of Eulimnadia texana Packard, 1871 (Branchiopoda: Spinicaudata) resistant to invasion? Implications for the maintenance of androdioecy. Journal of Crustacean Biology 38(6):812-817.

Ford, R. and S.C. Weeks. 2018. Intersexual conflict in androdioecious clam shrimp: Do androdioecious hermaphrodites evolve to avoid mating with males? Ethology 124:357-364. DOI:10.1111/eth.12738

Baldwin-Brown, J.G., S.C. Weeks and A.D. Long. 2018. A new standard for crustacean genomes: the highly contiguous, annotated genome assembly of the clam shrimp Eulimnadia texana reveals HOX gene order and identifies the sex chromosome. Genome Biology and Evolution 10:143-156. (Also noted inBioRxiv doi:

Roketenetz, L, S Donnelly, and SC Weeks. 2017. Phylogeographic characterization of genetic variation in the biological control agent milfoil weevil (Euhrychiopsis lecontei) throughout North America. American Midland Naturalist 178:260-274.

Calabrese, A, C McCullough, B Knott and SC Weeks. 2016. Environmental characteristics of ephemeral rock pools explain local abundances of the clam shrimp, Paralimnadia badia (Branchiopoda: Spinicaudata: Limnadiidae). Journal of the Royal Society of Western Australia 99:9-15.

Rogers, DC, N Rabet, and SC Weeks. 2016. Gondwanalimnadia (Branchiopoda: Spinicaudata) replacement name for Afrolimnadia Rogers, Rabet and Weeks, 2012 (Limnadiidae), junior homonym of Afrolimnadia Tasch, 1987 (Lioestheriidae). Journal of Crustacean Biology 36:105.

Reed, SK, RJ Duff, and SC Weeks. 2015. A systematic study of the genus Eulimnadia. Journal of Crustacean Biology 35:379-391.

Weeks, SC, C. Benvenuto, SK Reed, RJ Duff, and ZH Duan. 2014. A field test of a model for the stability of androdioecy in the freshwater shrimp, Eulimnadia texana. Journal of Evolutionary Biology 41:251-261.

Brown, B.P., T.I. Astrop and S.C. Weeks. 2014. Post-larval developmental dynamics of the Spinicaudatan (Crustacea: Branchiopoda) carapace. Journal of Crustacean Biology 34:611-617.

Weeks, SC, JS Brantner, TI Astrop, DW Ott and N Rabet. 2014. The evolution of hermaphroditism from dioecy in crustaceans: Selfing hermaphroditism described in a fourth Spinicaudatan genus. Evolutionary Biology 41:251-261. DOI 10.1007/s11692-013-9265-0.

Brantner, J. S. , D. W. Ott, R. J. Duff, L-O Sanoamuang, GP Simhachalam, KK Subhash Babu, and SC Weeks. 2013. Androdioecy and Hermaphroditism in Five Species of Clam shrimp (Crustacea: Branchiopoda: Spinicaudata) From India and Thailand. Invert. Biol. 132:27-37.

Brantner, J. S. , D. W. Ott, R. J. Duff, J.I. Orridge, J. R. Waldman and S.C. Weeks. 2013. Evidence of selfing hermaphroditism in the clam shrimp Cyzicus gynecia (Branchiopoda, Spinicaudata). J. Crust. Biol. 33:184-190.

Weeks, S.C. 2012. The role of androdioecy and gynodioecy in mediating evolutionary transitions between dioecy and hermaphroditism in the Animalia. Evolution 66:3670-3686.

Astrop, TI , LE Park, B Brown and SC Weeks. 2012. Sexual discrimination at work: Spinicaudatan ‘Clam Shrimp’ (Crustacea: Branchiopoda) as a model organism for the study of sexual system evolution. Palaeontologia Electronica 15.2.20A.

Rogers, DC, N Rabet, and SC Weeks. 2012. A revision of the extant genera of the Limnadiidae (Branchiopoda, Spinicaudata) J. Crust. Biol. 32:827-842.

Benvenuto, C, and SC Weeks. 2012. Intersexual conflict during mate guarding in an androdioecious crustacean. Behav. Ecol. 23:218–224.

Benvenuto C,and SC Weeks. 2011. Mate guarding behavior in clam shrimp: The influence of mating system on intersexual conflict. Behav. Ecol. Sociobiol. 65:1899-1907.

Weeks SC, TF Sanderson , BF Wallace, and B Bagatto. 2011. Behavioral cost of reproduction in a freshwater crustacean (Eulimnadia texana). Ethology 117:880-886.

Weeks SC, C Benvenuto , TF Sanderson, and RJ Duff. 2010. Sex Chromosome Evolution in the Clam Shrimp, Eulimnadia texana. J. Evol. Biol. 23:1100-1106.

Rogers DC, SC Weeks, and WR Hoeh. 2010. A new species of Eulimnadia (Crustacea; Branchiopoda; Diplostraca; Spinicaudata) from North America. Zootaxa 2413:61-68.

Weeks SC, EG Chapman , DC Rogers, DM Senyo, and WR Hoeh. 2009. Evolutionary transitions among dioecy, androdioecy and hermaphroditism in limnadiid clam shrimp (Branchiopoda: Spinicaudata). J. Evol. Biol. 22:1781-1799.

Benvenuto C, B Knott and SC Weeks. 2009. Mate guarding behavior in clam shrimp: A field approach. Behav. Ecol. 20:1125-1132.

Weeks SC, SK Reed , DW Ott, and F. Scanabissi. 2009. Inbreeding effects on sperm production in clam shrimp (Eulimnadia texana). Evol. Ecol. Res. 11:125-134.

Weeks SC. 2009. Can males successfully invade hermaphroditic populations of clam shrimp (Eulimnadia texana)? Current Science 96:98-102.

Weeks SC, TF Sanderson , M Zofkova, and B Knott. 2008. Breeding systems in the clam shrimp family Limnadiidae. Invertebrate Biology 127:336-349.

Brendonck L, DC Rogers, J Olesen, S Weeks, R Hoeh. 2008. Global diversity of large branchiopods (Crustacean; Branchiopoda) in freshwater. Hydrobiologia 595:167-176.

Weeks SC, and C Benvenuto. 2008. Mate guarding in the androdioecious clam shrimp Eulimnadia texana: male assessment of hermaphrodite receptivity. Ethology 114:64-74.

Duff RJ, C Benvenuto, T Branch, and SC Weeks. 2007. DNA extraction from cysts of the clam shrimp Eulimnadia texana (Crustacea, Branchiopoda, Spinicaudata). Journal of Crustacean Biology 27:154-157.

Weeks SC, M Zofkova, and B. Knott. 2006. Limnadiid Clam Shrimp Distribution in Australia (Crustacea: Branchiopoda: Spinicaudata). Journal of the Royal Society of Western Australia 89:155-161.

Weeks SC, SK Reed, M Cesari, and F Scanabissi. 2006. Production of intersexes and the evolution of androdioecy in the clam shrimp Eulimnadia texana (Crustacea, Branchiopoda, Spinicaudata). Invertebrate Reproductive Development 49:113-119.

Weeks SC, C Benvenuto, and SK Reed. 2006. When males and hermaphrodites coexist: A review of androdioecy in animals. Integrative and Comparative Biology 46:449-464.

Hoeh WR , ND Smallwood , DM Senyo, EG Chapman, and SC Weeks. 2006. Evaluating the monophyly of Eulimnadia and the Limnadiinae (Branchiopoda: Spinicaudata) using DNA sequences. Journal of Crustacean Biology 26:182-192.

Weeks SC, TF Sanderson, SK Reed, M Zofkova, B Knott, U Balaraman, G Pereira, DM Senyo, WR Hoeh. 2006. Ancient Androdioecy in the Freshwater Crustacean Eulimnadia . Proceedings of the Royal Society of London, Series B 273:725-734 .

Scanabissi F, M Cesari, SK Reed, SC Weeks. 2006. Ultrastructure of the male gonad and male gametogenesis in Eulimnadia texana (Packard, 1871)(Crustacea, Branchiopoda, Spinicaudata). Invertebrate Biology 125:117-124.

Weeks, SC, RT Posgai, M Cesari, and F Scanabissi. 2005. Androdioecy inferred in the clam shrimp Eulimnadia agassizii (Spinicaudata: Limnadiidae). Journal of Crustacean Biology 25:323-328.

Duff, RJ WR Hoeh, DG Cook and SC Weeks. 2004. Isolation and characterization of thirteen polymorphic microsatellite loci from the clam shrimp Eulimnadia texana (Crustacea:Spinicaudata). Molecular Ecology Notes 4:397-399.

Weeks, SC. 2004. Seven generations of inbreeding does not purge inbreeding depression in the clam shrimp, Eulimnadia texana. Journal of Evolutionary Biology 17:475-484.

Weeks, SC and RL Bernhardt. 2004. Maintenance of androdioecy in the freshwater shrimp, Eulimnadia texana: Field estimates of inbreeding depression and relative male survival. Evolutionary Ecology Research 6:227-242.

Weeks SC, CL Marquette, and E Latsch. 2004. Barriers to outcrossing success in the primarily self-fertilizing clam shrimp, Eulimnadia texana (Crustacea, Branchiopoda). Invertebrate Biology 123:146-155.

Weeks, SC and RJ Duff. 2002. A Genetic Comparison of Different Populations of Clam Shrimp in the genus Eulimnadia. Hydrobiologia 486:295-302.

Weeks, SC, V Marcus, R Salisbury, and D Ott. 2002. Cyst Development in the Conchostracan shrimp, Eulimnadia texana. Hydrobiologia 486:289-294.

Hollenbeck VG, N Zucker, W Gould and SC Weeks. 2002. Maintenance Of Androdioecy In The Freshwater Shrimp Eulimnadia texana: Sexual Encounter Rates And Outcrossing Success. Behav. Ecol. 13:561-570.

Naida Zucker, Gabriela A. Aguilar, Stephen C. Weeks,  and L. Garner McCandless. 2002. Variation in reproductive cycle between selfing and outcrossing hermaphrodites in an androdioecious desert shrimp. Invertebrate Biology 121:66-72.

Weeks, SC, J Hutchison, and N Zucker. 2001. Maintenance of androdioecy in the freshwater shrimp, Eulimnadia texana: Do hermaphrodites need males for complete fertilization? Evolutionary Ecology 15:205-221.

Gray, M and SC Weeks. 2001. A comparison of dietary use patterns of clonal and sexual fish (Poeciliidae: Poeciliopsis). Can. J. Fish. Aquat. Sci. 58:1313-1318.

Zucker, N, B Stafki and SC Weeks. 2001. Maintenance of androdioecy in the freshwater shrimp, Eulimnadia texana: Relative longevity of males to hermaphrodites. Can. J. Zool. 79:393-401.

Weeks, SC, BR Crosser, and MM Gray. 2001. Relative fitness of two hermaphroditic mating types in the androdioecious clam shrimp, Eulimnadia texana. Journal of Evolutionary Biology 14:83-94.

Medland, VL, N Zucker, and SC Weeks. 2000. Maintenance of androdioecy in the freshwater shrimp, Eulimnadia texana Parker: Encounters between males and hermaphrodites are not random. Ethology 106:839-848.

Weeks, SC, BR Crosser, B Bennett, MM Gray, and N Zucker. 2000. Maintenance of androdioecy in the freshwater shrimp, Eulimnadia texana: Variation in inbreeding depression between two populations.  Evolution 54:878-887.

Weeks, SC, BR Crosser, MM Gray, JA Matweyou, and N. Zucker. 2000. Is there Sperm Storage in the Clam Shrimp Eulimnadia texana? Invertebrate Biology 119:215-221.

Weeks, SC, and N Zucker. 1999. Rates of Inbreeding in the Clam Shrimp Eulimnadia texana. Can. J. Zool. 77:1402-1408.

Stockwell, CA and SC Weeks. 1999. Translocations and altered evolutionary trajectories in recently established populations of the western mosquitofish (Gambusia affinis) Animal Conservation 2:103-110.

Weeks, SC, V Marcus, and B Crosser. 1999. Inbreeding Depression in a Self-Compatible, Androdioecious Crustacean, Eulimnadia texana. Evolution 53:472-483.

Belk, D, G Mura and SC Weeks. 1998. Untangling confusion between Eubranchipus vernalis and Eubranchipus neglectus (Branchiopoda, Anostraca). Journal of Crustacean Biology 18:147-152.

Weeks, SC and V Marcus. 1997. A survey of the branchiopod crustaceans of Ohio. Ohio Journal of Science 97:86-89.

Marcus, V and SC Weeks. 1997. The effects of pond duration on the life history traits of an ephemeral pond crustacean, Eulimnadia texana. Hydrobiologia, 359:213-221.

Weeks, SC and GK Meffe. 1997. “Grandfather Effects” on offspring size in the Eastern Mosquitofish, Gambusia holbrooki. Copeia 1997:869-874.

Weeks, SC, V Marcus and S Alvarez. 1997. Notes on the life history of the clam shrimp Eulimnadia texana. Hydrobiologia 359:191-197.

Weeks SC. 1996. A reevaluation of Red Queen model for the maintenance of sex in a clonal-sexual fish complex (Poeciliidae: Poeciliopsis). Canadian Journal of Fisheries and Aquatic Sciences 53:1157-1164.

Weeks SC. 1996. The hidden cost of reproduction: reduced energetic intake caused by spatial constraints in the body cavity. Oikos 75:345-349.

Weeks SC and GK Meffe. 1996. Quantitative genetic and optimality analyses of life-history plasticity in the eastern mosquitofish, Gambusia holbrooki. Evolution 50:1358-1365.

Meffe, G.K., S.C. Weeks, P. Mulvey, and K.L. Kandl. 1995. Genetic differences in thermal tolerance of mosquitofish from ambient and thermally elevated ponds. Canadian Journal of Fisheries and Aquatic Sciences 52:2704-2711.

Weeks, S.C. 1995. Comparisons of life-history traits between clonal and sexual fish (Poeciliopsis: Poeciliidae) raised in monoculture and mixed treatments. Evolutionary Ecology 9:258-274.

Heulett ST, SC Weeks and GK Meffe. 1995. Lipid dynamics and growth relative to resource level in juvenile eastern mosquitofish (Gambusia holbrooki: Poeciliidae). Copeia 1995:97-104.

Weeks SC and OE Gaggiotti. 1993. Patterns of offspring size at birth in clonal and sexual strains of Poeciliopsis (Poeciliidae). Copeia 1993:1003-1009.

Weeks SC. 1993. The effects of recurrent clonal formation on clonal invasion patterns and sexual persistence: A Monte Carlo simulation of the frozen niche-variation model. The American Naturalist 141:409-427.

Sassaman C and SC Weeks. 1993. The genetic mechanism of sex determination in the Conchostracan shrimp Eulimnadia texana. The American Naturalist 141:314-328.

Weeks SC. 1993. Phenotypic plasticity of life-history traits in clonal and sexual fish (Poeciliopsis) at high and low densities. Oecologia 93:307-314.

Connor A, KM Weiss and SC Weeks. 1993. Evolutionary models of quantitative disease risk factors. Human Biology 65:917-940.

Weeks SC, OE Gaggiotti, R Schenck, K Spindler, and RC Vrijenhoek. 1992. Feeding behavior in sexual and clonal strains of Poeciliopsis. Behavioral Ecology and Sociobiology 30:1-6.

Weeks SC and JM Quattro. 1991. Life-history plasticity under resource stress in a clonal fish (Poeciliidae: Poeciliopsis). Journal of Fish Biology 39:485-494.

Quattro JM and SC Weeks. 1991. Correlations of egg size and energetic content in sexual and clonal biotypes of Poeciliopsis. Journal of Fish Biology 38:331-334.

Weeks SC and C Sassaman . 1990. Competition in phenotypically variable and uniform populations of the tadpole shrimp Triops longicaudatus (Notostraca: Triopsidae). Oecologia82:552-559.

Weeks SC. 1990. Life-history variation under varying degrees of intraspecific competition in the tadpole shrimp Triops longicaudatus (LeConte). Journal of Crustacean Biology 10:498-503.

Wetherington JD, SC Weeks, KE Kotora, and RC Vrijenhoek. 1989. Genotypic and environmental components of variation in growth and reproduction of fish hemiclones(Poeciliopsis: Poeciliidae). Evolution 43:635-645.

Book Chapters

Benvenuto, C and SC Weeks. 2020. Hermaphroditism and Gonochorism. Pp. 197-241 In: Cothran, R. and M. Thiel (eds), The Natural History of the Crustacea. Vol. 6. Reproductive Biology. Oxford University Press, New York, NY.

Benvenuto, C., B Knott, and SC Weeks. 2015. Crustaceans of extreme environments. Pp. 379-417 in: The Natural History of the Crustacea, Vol. 2: Lifestyles and Feeding Biology, Thiel, M. and L. Watling, eds. Oxford University Press, New York, NY.

Colburn EA, SC Weeks, SK Reed. 2007. Diversity and Ecology of Vernal Pool Invertebrates. pp. 105 – 126 in: Science and Conservation of Vernal Pools in Northeastern North America, Calhoun A and P DeMaynadier, eds. CRC Press.

Wood JW, SC Weeks, GR Bentley and KM Weiss. 1994. Human population biology and the evolution of aging. Pp. 19-75 in: Crews DE and RM Garruto (eds.) Biological Anthropology and Aging. Perspectives on Human Variation Over the Life Span. Oxford University Press, New York, NY.


Non-referred Publications

Weeks SC. 2006. Under the flashlight: Clam shrimp. Vernalis 9:1-6.

Baldwin-Brown, J.G., S.C. Weeks and A.D. Long. 2017. A new standard for crustacean genomes: the highly contiguous, annotated genome assembly of the clam shrimp Eulimnadia texana reveals HOX gene order and identifies the sex chromosome. BioRxiv doi: